Everard Millais. He offered me every facility. The Basset Hound records referring to his own and other breeds had been carefully kept, and the Stud Book he lent me contained accounts of nearly moo animals, of which I was able to utilise 817. All were descended from parents of known colours ; in 567 of them the colours of all four grandparents were also known. Wherever the printed Stud Book was deficient, Sir Everard Millais supplied the want in MS from the original records. My inquiry was into the heredity of two alternative colours, one containing no black, the other containing it; their technical names were lemon-white and tri-colour (black, lemon, white) respectively. I was assured that no difficulty was felt in determining the category to which each individual belonged. These data were fully discussed in a memoir, published (1897) in the Proceedings of the Royal Society [i 39], on what is now termed the " Ancestral Law," namely, that the average contribution of each parent is 4, of each grandparent ia, and so on. Or, in other words, that of the two parents taken together is 1, of the four grandparents together 1, and so on. My data were not as numerous as is desirable, still the results were closely congruous, and seem to be a near approximation to the truth. The conclusions have been much discussed and criticised, and they have been modified by Professor Karl Pearson ; but they have not been seriously shaken, so far as I know.
